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He spent most of his professional career with Villarreal after signing as a 19-year-old, going on to appear in 249 competitive matches and score seven goals during seven La Liga seasons. In 2017, he joined Milan.

On 30 May 2017, eight days after passing his medical, Musacchio moved to Italian club A.C. Milan after signing a four-year contract. He contributed with 15 Serie A appearances in his first season, with the subsequent qualification to the UEFA Europa League after a sixth-place finish.

In 2017, Gianni De Biasi asked Musacchio if he would be interested in playing for Albania of which he was the manager, but the player declined as he wished to represent his country of birth.

Musacchio was first called by the Argentina national team in May 2011, as manager Sergio Batista only selected players under 25 for friendlies with Nigeria and Poland. He made his debut against the former on 2 June, in a 1–4 loss.

On 13 February 2010, shortly after ceasing to be a non-EU player, Musacchio made his La Liga debut by playing the last 15 minutes in a 2–1 home win against Athletic Bilbao. From that moment onwards he was permanently promoted to Villarreal's first team, successively surpassing more experienced Iván Marcano and Gonzalo Rodríguez – the latter his compatriot – in the defensive pecking order.

In August 2009, Musacchio moved to Villarreal CF in Spain, initially being assigned to the B-team in the second division. He made his official debut on 5 September in a 1–3 loss at Córdoba CF as the reserves overachieved in their first season ever in that tier, eventually finishing seventh.

A product of Club Atlético River Plate's youth system, Musacchio was born in Rosario, Santa Fe, and he made his first-team debut as a 16-year-old during the 2006–07 season, making four appearances. He was part of the squad that won the Clausura in the following year, but did not feature in any games.

Mateo Pablo Musacchio (Spanish: [maˈte.o muˈsakjo] ; born 26 August 1990) is an Argentine footballer who plays as a central defender for Italian club A.C. Milan.